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An application layer is an abstraction layer that specifies the shared protocols and interface methods used by hosts in a communications network. The application layer abstraction is used in both of the standard models of computer networking; the Internet Protocol Suite (TCP/IP) and the Open Systems Interconnection model (OSI model).

In TCP/IP, the application layer contains the communications protocols and interface methods used in process-to-process communications across an Internet Protocol (IP) computer network. The application layer only standardizes communication and depends upon the underlying transport layer protocols to establish host-to-host data transfer channels and manage the data exchange in a client-server or peer-to-peer networking model. Though the TCP/IP application layer does not describe specific rules or data formats that applications must consider when communicating, the original specification.

The OSI model defines the application layer as the user interface responsible for displaying received information to the user. In contrast, the Internet Protocol model does not concern itself with such detail. OSI also explicitly distinguishes additional functionality below the application layer, but above the transport layer at two additional levels; the session layer and the presentation layer. OSI also specifies a strict modular separation of functionality at these layers and provides protocol implementations for each layer.